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High-profile newcomer Storm Stacey led the Superstock speeds for much of their slightly curtailed session but ultimately ended up third on the leaderboard with 107.870mph.
Australian Josh Brookes came out on top on 109.411, with Harrison again to the fore in second place at 108.814.
John McGuinness, Dominic Herbertson and Hickman were next in the order.
Both Brookes and Harrison are chasing their first-ever wins at the North West.
McWilliams was 7.7 seconds faster than his nearest rival in producing the best speed of the Supertwins/Sportbike session on 105.163mph on drying roads on his Yamaha R7.
Jordan, winner of last year's second Supertwins event, was his closest challenger at 102.590, with Seeley third fastest at 101.502, both riding Aprilias.
Record 29-time winner Seeley is making his first appearance in the Supertwins category.
Michael Dunlop endured a disappointing day as he failed to feature in the upper echelons of the Superbike leaderboard and has yet to qualify after making a last-minute decision to switch from his Ducati V4 to a Honda Fireblade in the Superbike class.
The Ballymoney rider was 12th in the Supersports and back in 25th position in the Superstock outing.

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